what is the final product and main goal of photosynthesis

What Is The End Product Of Photosynthesis?

Humans and most other animals need certain things to survive. Oxygen is one of them, and the carbohydrate glucose is another. Fortunately for them, plants (and certain bacteria and algae) produce both of these as the result of a complex process known as photosynthesis.

The Formula

The formula associated with the process of photosynthesis is

6H 2 O + 6CO 2 = C 6 H 12 O 6 + 6O 2 .

This formula tells you is that six molecules of water plus six molecules of carbon dioxide will produce one molecule of glucose plus six molecules of oxygen. This entire process goes through two distinct stages before it is completed. The first stage is a light-dependent process and the second stage is a light-independent process.

Light Dependent

In the light-dependent process, the electrons of the chloroplasts (special organelles used to carry out photosynthesis) are excited into a higher energy state when they are bombarded with light. These excited electrons cause a series of reactions that produce adenosine triphosphate (ATP) and nicotinamide adenine dinucleotide phosphate (NADPH). ATP and NADPH are then used to make carbon bonds in the light-independent process. Water molecules present in the light-dependent process are split. Their oxygen molecules are released into the atmosphere.

Light Independent

Recall the splitting of the water molecules in the light-dependent process that released oxygen molecules into the atmosphere. Since water is H 2 0, there is still a hydrogen atom remaining. This hydrogen atom is used in the light-independent process when plants take carbon dioxide from the atmosphere. The carbon dioxide and hydrogen become bound together through a process called carbon fixation, which forms a non-specific carbohydrate.

Photophosphorylation

Photophosphorylation is the process by which light energy produces NADPH. Special pigments found in the plant's cells known as chlorophyll make this process possible. The two main types of chlorophyll are chlorophyll A and chlorophyll B. In simple terms, the electrons of water molecules present in chlorophyll B become excited by the presence of light. Chlorophyll B takes one of these excited electrons splitting the H 2 O molecule into H + and O -2 . O -2 is converted into O 2 and released into the atmosphere. The excited electron is attached to a primary electron receptor, and through a series of complex reactions forms NADPH. NADPH is the energy carrier used in carbon fixation.

The Calvin Cycle

Plants produce glucose in a process known as the Calvin cycle. The carbon dioxide captured in the light-independent process is processed in this cycle. For every six molecules of carbon dioxide captured and put into the cycle, one molecule of glucose is produced. The chemical that captures the carbon dioxide for use in the Calvin cycle is ribulose biphosphate.

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Cite This Article

Boles, Kelley. "What Is The End Product Of Photosynthesis?" sciencing.com , https://www.sciencing.com/end-product-photosynthesis-6570858/. 13 April 2018.

Boles, Kelley. (2018, April 13). What Is The End Product Of Photosynthesis?. sciencing.com . Retrieved from https://www.sciencing.com/end-product-photosynthesis-6570858/

Boles, Kelley. What Is The End Product Of Photosynthesis? last modified March 24, 2022. https://www.sciencing.com/end-product-photosynthesis-6570858/

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What Are the Products of Photosynthesis?

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What are the products of photosynthesis? First, let's define this process: Photosynthesis is the name given to the set of chemical reactions performed by plants to convert energy from the sun into chemical energy in the form of sugar. Specifically, plants use energy from sunlight to react to carbon dioxide and water to produce sugar (glucose) and oxygen, the products of photosynthesis.

Many reactions occur, but the overall chemical reaction for photosynthesis is:

• 6 CO 2 + 6 H 2 O + light → C 6 H 12 O 6 + 6 O 2
• Carbon Dioxide + Water + Light yields Glucose + Oxygen

In a plant, the carbon dioxide enters via leaf stomates by diffusion. Water is absorbed through the roots and is transported to leaves through the xylem. Solar energy is absorbed by chlorophyll in the leaves. The reactions of photosynthesis occur in the chloroplasts of plants. In photosynthetic bacteria, the process takes place where chlorophyll or a related pigment is embedded in the plasma membrane. The oxygen and water produced in photosynthesis exit through the stomata.

Key Takeaways

• In photosynthesis, energy from light is used to convert carbon dioxide and water into glucose and oxygen.
• For 6 carbon dioxide and 6 water molecules, 1 glucose molecule and 6 oxygen molecules are produced.

Actually, plants reserve very little of the glucose for immediate use. Glucose molecules are combined by dehydration synthesis to form cellulose, which is used as a structural material. Dehydration synthesis is also used to convert glucose to starch, which plants use to store energy.

Intermediate Products of Photosynthesis

The overall chemical equation is a summary of a series of chemical reactions. These reactions occur in two stages. The light reactions require light (as you might imagine), while the dark reactions are controlled by enzymes. They don't require darkness to occur—they simply don't depend on light.

The light reactions absorb light and harness the energy to power electron transfers. Most photosynthetic organisms capture visible light, although there are some that use infrared light. Products of photosynthesis are adenosine triphosphate ( ATP ) and reduced nicotinamide adenine dinucleotide phosphate (NADPH). In plant cells, the light-dependent reactions occur in the chloroplast thylakoid membrane. The overall reaction for the light-dependent reactions is:

• 2 H 2 O + 2 NADP +  + 3 ADP + 3 P i  + light → 2 NADPH + 2 H +  + 3 ATP + O 2

In the dark stage, ATP and NADPH ultimately reduce carbon dioxide and other molecules. Carbon dioxide from the air is "fixed" into a biologically usable form, glucose . In plants, algae, and cyanobacteria, the dark reactions are termed the Calvin cycle. Bacteria may use different reactions, including a reverse Krebs cycle . The overall reaction for the light-independent reaction of a plant (Calvin cycle) is:

• 3 CO 2  + 9 ATP + 6 NADPH + 6 H +  → C 3 H 6 O 3 -phosphate + 9 ADP + 8 P i  + 6 NADP +  + 3 H 2 O

During carbon fixation, the three-carbon product of the Calvin cycle is converted into the final carbohydrate product.

Factors That Affect the Rate of Photosynthesis

Like any chemical reaction, the availability of the reactants determines the amount of products of photosynthesis that can be made. Limiting the availability of carbon dioxide or water slows the production of glucose and oxygen . Also, the rate of the reactions is affected by temperature and the availability of minerals that may be needed in the intermediate reactions.

The overall health of the plant (or other photosynthetic organism) also plays a role. The rate of metabolic reactions is determined in part by the maturity of the organism and whether it's flowering or bearing fruit.

What Is Not a Product of Photosynthesis?

If you're asked about this process on a test, you may be asked to identify the products of photosynthesis . That's pretty easy, right? Another form of the question is to ask what is not a product of photosynthesis. Unfortunately, this won't be an open-ended question, which you could easily answer with "iron" or "a car" or "your mom." Usually this is a multiple choice question, listing molecules which are reactants or products of photosynthesis. The answer is any choice except glucose or oxygen. The question may also be phrased to answer what is not a product of the light reactions or the dark reactions. So, it's a good idea to know the overall reactants and products for the photosynthesis general equation, the light reactions, and the dark reactions.

• Bidlack, J.E.; Stern, K.R.; Jansky, S. (2003). Introductory Plant Biology . New York: McGraw-Hill. ISBN 978-0-07-290941-8.
• Blankenship, R.E. (2014). Molecular Mechanisms of Photosynthesis (2nd ed.). John Wiley & Sons. ISBN 978-1-4051-8975-0.
• Reece J.B., et al. (2013). Campbell Biology . Benjamin Cummings. ISBN 978-0-321-77565-8.
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What is the end product of photosynthesis?

Photosynthesis: photosynthesis is the process by which plants produce their own food from raw resources such as sunlight, chlorophyll, water, and carbon dioxide. it is the primary source of all food on earth. it is also responsible for the release of oxygen into the atmosphere by green plants. the end product of photosynthesis: glucose and oxygen are the final products of photosynthesis..

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Photosynthesis

1. Photosynthesis is the process plants use to make their own food.

Like all living things, plants need energy to carry out the processes that keep them alive. They get this energy from food. Humans and most other animals are heterotrophs, meaning we have to consume other organisms—plants, other animals, or some combination of the two—for food. However, plants are autotrophs, meaning they create their own food.

Plants use sunlight to convert water and carbon dioxide into glucose and oxygen in a process called photosynthesis . In biology, this information is often expressed using a chemical equation .

Chemical equations typically show the molecules that enter the reaction (the reactants ) to the left and the molecules that result from the reaction (the products ) to the right, separated by an arrow that indicates a reaction taking place.

[Reactants] → [Products]

You can think of the reactants as the ingredients for preparing a meal and the products as the different dishes in that meal.

With that in mind, let’s take a look at the chemical equation for photosynthesis:

Sunlight + 6 CO 2 + 6 H 2 O → C 6 H 12 O 6 + 6 O 2 CO 2 = carbon dioxide H 2 O = water C 6 H 12 O 6 = glucose O 2 = oxygen * Sometimes, you’ll see sunlight, or a symbol indicating the sun, over the arrow in the equation.

Therefore, to produce one molecule of glucose (and 6 molecules of oxygen gas), a plant needs 6 molecules of carbon dioxide and 6 molecules of water.

2. The reactants of photosynthesis are carbon dioxide and water.

We’ve established that plants need carbon dioxide (CO 2 ) and water (H 2 O) to produce their food, but where do these reactants come from and how do they get where they need to go inside the plant?

Plants take in carbon dioxide from the air through small openings in their leaves called stomata. Some plants (most monocots) have stomata on both sides of their leaves, and others (dicots and a few monocots) only have stomata on the underside, or lower epidermis.

Plants get water from the soil surrounding their roots, and water gets to the leaves by traveling through the xylem, part of the plant’s vascular system. In leaves, the xylem and phloem are contained in the vascular bundle.

Once inside the leaf, the carbon dioxide and water molecules move into the cells of the mesophyll, the layer of ground tissue between the upper and lower epidermis. Within these cells, organelles called chloroplasts use the carbon dioxide and water to carry out photosynthesis.

3. Light energy from the sun initiates photosynthesis in the chloroplasts of plant cells.

Plant cells have special organelles called chloroplasts, which serve as the sites for the reactions that make up photosynthesis. Their thylakoid membranes contain a pigment called chlorophyll, which absorbs photons (light energy) from the sun, initiating the light-dependent reactions that take place within the thylakoids.

During these reactions, water molecules (H 2 O) are broken down. NADPH and ATP—high energy molecules that power the production of glucose—are produced during the light-dependent reactions, as well. Electrons and hydrogen ions from the water are used to build NADPH. Hydrogen ions also power the conversion of ADP to ATP.

4. The products of photosynthesis are glucose and oxygen.

Did you know that oxygen is actually a waste product of photosynthesis? Although the hydrogen atoms from the water molecules are used in the photosynthesis reactions, the oxygen molecules are released as oxygen gas (O 2 ). (This is good news for organisms like humans and plants that use oxygen to carry out cellular respiration!) Oxygen passes out of the leaves through the stomata.

The light-independent reactions of photosynthesis—also known as the Calvin cycle—use enzymes in the stroma, along with the energy-carrying molecules (ATP and NADPH) from the light-dependent reactions, to break down carbon dioxide molecules (CO 2 ) into a form that is used to build glucose.The mitochondria in the plant’s cells use cellular respiration to break glucose down into a usable form of energy (ATP), which fuels all the plant’s activities.

After the light-independent reactions, glucose is often made into larger sugars like sucrose or carbohydrates like starch or cellulose. Sugars leave the leaf through the phloem and can travel to the roots for storage or to other parts of the plant, where they’re used as energy to fuel the plant’s activities.

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8. Photosynthesis

Overview of photosynthesis, learning objectives.

By the end of this section, you will be able to do the following:

• Explain the significance of photosynthesis to other living organisms
• Describe the main structures involved in photosynthesis
• Identify the substrates and products of photosynthesis

Photosynthesis is essential to all life on earth; both plants and animals depend on it. It is the only biological process that can capture energy that originates from sunlight and converts it into chemical compounds (carbohydrates) that every organism uses to power its metabolism. It is also a source of oxygen necessary for many living organisms. In brief, the energy of sunlight is “captured” to energize electrons, whose energy is then stored in the covalent bonds of sugar molecules. How long lasting and stable are those covalent bonds? The energy extracted today by the burning of coal and petroleum products represents sunlight energy captured and stored by photosynthesis 350 to 200 million years ago during the Carboniferous Period.

Plants, algae, and a group of bacteria called cyanobacteria are the only organisms capable of performing photosynthesis ( (Figure) ). Because they use light to manufacture their own food, they are called photoautotrophs (literally, “self-feeders using light”). Other organisms, such as animals, fungi, and most other bacteria, are termed heterotrophs (“other feeders”), because they must rely on the sugars produced by photosynthetic organisms for their energy needs. A third very interesting group of bacteria synthesize sugars, not by using sunlight’s energy, but by extracting energy from inorganic chemical compounds. For this reason, they are referred to as chemoautotrophs.

The importance of photosynthesis is not just that it can capture sunlight’s energy. After all, a lizard sunning itself on a cold day can use the sun’s energy to warm up in a process called behavioral thermoregulation . In contrast, photosynthesis is vital because it evolved as a way to store the energy from solar radiation (the “photo-” part) to energy in the carbon-carbon bonds of carbohydrate molecules (the “-synthesis” part). Those carbohydrates are the energy source that heterotrophs use to power the synthesis of ATP via respiration. Therefore, photosynthesis powers 99 percent of Earth’s ecosystems. When a top predator, such as a wolf, preys on a deer ( (Figure) ), the wolf is at the end of an energy path that went from nuclear reactions on the surface of the sun, to visible light, to photosynthesis, to vegetation, to deer, and finally to the wolf.

Main Structures and Summary of Photosynthesis

Photosynthesis is a multi-step process that requires specific wavelengths of visible sunlight, carbon dioxide (which is low in energy), and water as substrates ( (Figure) ). After the process is complete, it releases oxygen and produces glyceraldehyde-3-phosphate (GA3P), as well as simple carbohydrate molecules (high in energy) that can then be converted into glucose, sucrose, or any of dozens of other sugar molecules. These sugar molecules contain energy and the energized carbon that all living things need to survive.

The following is the chemical equation for photosynthesis ( (Figure) ):

Although the equation looks simple, the many steps that take place during photosynthesis are actually quite complex. Before learning the details of how photoautotrophs turn sunlight into food, it is important to become familiar with the structures involved.

Basic Photosynthetic Structures

In plants, photosynthesis generally takes place in leaves, which consist of several layers of cells. The process of photosynthesis occurs in a middle layer called the mesophyll. The gas exchange of carbon dioxide and oxygen occurs through small, regulated openings called stomata (singular: stoma), which also play roles in the regulation of gas exchange and water balance. The stomata are typically located on the underside of the leaf, which helps to minimize water loss due to high temperatures on the upper surface of the leaf. Each stoma is flanked by guard cells that regulate the opening and closing of the stomata by swelling or shrinking in response to osmotic changes.

In all autotrophic eukaryotes, photosynthesis takes place inside an organelle called a chloroplast. For plants, chloroplast-containing cells exist mostly in the mesophyll. Chloroplasts have a double membrane envelope (composed of an outer membrane and an inner membrane), and are ancestrally derived from ancient free-living cyanobacteria. Within the chloroplast are stacked, disc-shaped structures called thylakoids. Embedded in the thylakoid membrane is chlorophyll, a pigment (molecule that absorbs light) responsible for the initial interaction between light and plant material, and numerous proteins that make up the electron transport chain. The thylakoid membrane encloses an internal space called the thylakoid lumen. As shown in (Figure) , a stack of thylakoids is called a granum, and the liquid-filled space surrounding the granum is called stroma or “bed” (not to be confused with stoma or “mouth,” an opening on the leaf epidermis).

Art Connection

On a hot, dry day, the guard cells of plants close their stomata to conserve water. What impact will this have on photosynthesis?

The Two Parts of Photosynthesis

Photosynthesis takes place in two sequential stages: the light-dependent reactions and the light-independent reactions. In the light-dependent reactions, energy from sunlight is absorbed by chlorophyll and that energy is converted into stored chemical energy. In the light-independent reactions, the chemical energy harvested during the light-dependent reactions drives the assembly of sugar molecules from carbon dioxide. Therefore, although the light-independent reactions do not use light as a reactant, they require the products of the light-dependent reactions to function. In addition, however, several enzymes of the light-independent reactions are activated by light. The light-dependent reactions utilize certain molecules to temporarily store the energy: These are referred to as energy carriers . The energy carriers that move energy from light-dependent reactions to light-independent reactions can be thought of as “full” because they are rich in energy. After the energy is released, the “empty” energy carriers return to the light-dependent reaction to obtain more energy. (Figure) illustrates the components inside the chloroplast where the light-dependent and light-independent reactions take place.

Link to Learning

Click the link to learn more about photosynthesis.

Everyday Connection

Photosynthesis at the Grocery Store

Major grocery stores in the United States are organized into departments, such as dairy, meats, produce, bread, cereals, and so forth. Each aisle ( (Figure) ) contains hundreds, if not thousands, of different products for customers to buy and consume.

Although there is a large variety, each item ultimately can be linked back to photosynthesis. Meats and dairy link, because the animals were fed plant-based foods. The breads, cereals, and pastas come largely from starchy grains, which are the seeds of photosynthesis-dependent plants. What about desserts and drinks? All of these products contain sugar—sucrose is a plant product, a disaccharide, a carbohydrate molecule, which is built directly from photosynthesis. Moreover, many items are less obviously derived from plants: For instance, paper goods are generally plant products, and many plastics (abundant as products and packaging) are derived from “algae” (unicellular plant-like organisms, and cyanobacteria). Virtually every spice and flavoring in the spice aisle was produced by a plant as a leaf, root, bark, flower, fruit, or stem. Ultimately, photosynthesis connects to every meal and every food a person consumes.

Section Summary

The process of photosynthesis transformed life on Earth. By harnessing energy from the sun, the evolution of photosynthesis allowed living things access to enormous amounts of energy. Because of photosynthesis, living things gained access to sufficient energy that allowed them to build new structures and achieve the biodiversity evident today.

Only certain organisms (photoautotrophs), can perform photosynthesis; they require the presence of chlorophyll, a specialized pigment that absorbs certain wavelengths of the visible spectrum and can capture energy from sunlight. Photosynthesis uses carbon dioxide and water to assemble carbohydrate molecules and release oxygen as a byproduct into the atmosphere. Eukaryotic autotrophs, such as plants and algae, have organelles called chloroplasts in which photosynthesis takes place, and starch accumulates. In prokaryotes, such as cyanobacteria, the process is less localized and occurs within folded membranes, extensions of the plasma membrane, and in the cytoplasm.

Art Connections

(Figure) On a hot, dry day, plants close their stomata to conserve water. What impact will this have on photosynthesis?

(Figure) Levels of carbon dioxide (a necessary photosynthetic substrate) will immediately fall. As a result, the rate of photosynthesis will be inhibited.

Review Questions

Which of the following components is not used by both plants and cyanobacteria to carry out photosynthesis?

• chloroplasts
• chlorophyll
• carbon dioxide

What two main products result from photosynthesis?

• oxygen and carbon dioxide
• chlorophyll and oxygen
• sugars/carbohydrates and oxygen
• sugars/carbohydrates and carbon dioxide

In which compartment of the plant cell do the light-independent reactions of photosynthesis take place?

• outer membrane

Which statement about thylakoids in eukaryotes is not correct?

• Thylakoids are assembled into stacks.
• Thylakoids exist as a maze of folded membranes.
• The space surrounding thylakoids is called stroma.
• Thylakoids contain chlorophyll.

Predict the end result if a chloroplast’s light-independent enzymes developed a mutation that prevented them from activating in response to light.

• GA3P accumulation
• ATP and NADPH accumulation
• Water accumulation
• Carbon dioxide depletion

Show Solution

How are the NADPH and GA3P molecules made during photosynthesis similar?

• They are both end products of photosynthesis.
• They are both substrates for photosynthesis.
• They are both produced from carbon dioxide.
• They both store energy in chemical bonds.

Free Response

What is the overall outcome of the light reactions in photosynthesis?

The outcome of light reactions in photosynthesis is the conversion of solar energy into chemical energy that the chloroplasts can use to do work (mostly anabolic production of carbohydrates from carbon dioxide).

Why are carnivores, such as lions, dependent on photosynthesis to survive?

Because lions eat animals that eat plants.

Why are energy carriers thought of as either “full” or “empty”?

The energy carriers that move from the light-dependent reaction to the light-independent one are “full” because they bring energy. After the energy is released, the “empty” energy carriers return to the light-dependent reaction to obtain more energy. There is not much actual movement involved. Both ATP and NADPH are produced in the stroma where they are also used and reconverted into ADP, Pi, and NADP+.

Describe how the grey wolf population would be impacted by a volcanic eruption that spewed a dense ash cloud that blocked sunlight in a section of Yellowstone National Park.

The grey wolves are apex predators in their food web, meaning they consume smaller prey animals and are not the prey of any other animal. Blocking sunlight would prevent the plants at the bottom of the food web from performing photosynthesis. This would kill many of the plants, reducing the food sources available to smaller animals in Yellowstone. A smaller prey animal population means that fewer wolves can survive in the area, and the population of grey wolves will decrease.

How does the closing of the stomata limit photosynthesis?

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What Are the Products of Photosynthesis?

Photosynthesis is a set of chemical reactions that plants and other organisms use to make chemical energy in the form of sugar. Like any chemical reaction, photosynthesis has reactants and products . Overall, the reactants of photosynthesis are carbon dioxide and water, while the products of photosynthesis are oxygen and glucose (a sugar).

Here’s a closer look at the products of photosynthesis and the balanced equation for the reaction.

The reactants for photosynthesis are carbon dioxide and water, while the products are the sugar glucose and oxygen.

Balanced Chemical Equation for Photosynthesis

Photosynthesis actually involves many chemical reactions, but the net balanced equation is that six moles of carbon dioxide react with six moles of water to produce one mole of glucose and six moles of oxygen. Light from the Sun provides the activation energy for the reaction. Sometimes light is listed in the balanced equation as a reactant, but it’s usually omitted.

6 CO 2  + 6 H 2 O → C 6 H 12 O 6  + 6 O 2

Carbon Dioxide + Water + Light → Glucose + Oxygen

Closer Look at the Products of Photosynthesis

Photosynthesis occurs in a series of steps that are classified as light-dependent reactions and light-independent reactions. Adding up the reactants and products of these reactions gives the overall equation for photosynthesis, but it’s good to know the inputs and outputs for each stage.

Light-Dependent Reactions

The light-dependent reactions or light reactions absorb certain wavelengths of light to make adenosine triphosphate (ATP) and reduced nicotinamide adenine dinucleotide phosphate (NADPH). The light reactions occur in the chloroplast thylakoid membrane. The overall balanced equation for the light-dependent reactions is:

2 H 2 O + 2 NADP +  + 3 ADP + 3 P i  + light → 2 NADPH + 2 H +  + 3 ATP + O 2

Light-Independent Reactions

While the light reactions use water, the light-independent reactions use carbon dioxide. The light-independent reactions are also called the dark reactions. These reactions do not require darkness, but they don’t depend on light to proceed. In plants, algae, and cyanobacteria, the dark reactions are called the Calvin cycle. Bacteria use different reactions, including the reverse Krebs cycle.

The overall balanced equation for the light-independent reactions (Calvin cycle) in plants is:

3 CO 2  + 9 ATP + 6 NADPH + 6 H +  → C 3 H 6 O 3 -phosphate + 9 ADP + 8 P i  + 6 NADP +  + 3 H 2 O

Finally, the three-carbon product from the Calvin cycle becomes glucose during the process of carbon fixation.

Other Products of Photosynthesis

Glucose is the direct product of photosynthesis, but plants turn most of the sugar into other compounds. These are indirect products. Linking glucose units forms starch and cellulose. Cellulose is a structural material. Plants store starch or link it to fructose (another sugar) to form sucrose (table sugar).

What Is Not a Product of Photosynthesis?

On an exam, you may need to identify which chemical is not a product of photosynthesis. For the overall process, choose any answer except “glucose” or “oxygen.” It’s good to know the overall reactants and products of the light reactions and dark reactions, in case you’re asked about them. The products of the light reactions are ATP , NADPH, protons, and oxygen. The products of the dark reactions are C 3 H 6 O 3 -phosphate, ADP, inorganic phosphate, NADP + , and water.

Where Does Photosynthesis Occur?

In addition to knowing the reactants and products of photosynthesis, you may need to know where photosynthesis occurs in different organisms.

• In plants, photosynthesis occurs in organelles called chloroplasts. Photosynthetic protists also contain chloroplasts. Leaves contain the highest concentration of chloroplasts in plants. Plants obtain carbon dioxide via diffusion through leaf stomata. Water comes from the roots and travels to the leaves via the xylem . Chlorophyll in chloroplasts absorbs solar energy. Oxygen from photosynthesis exits the plant via leaf stomata.
• Photosynthesis occurs in photosynthetic bacteria in the plasma membrane. Chlorophyll or related pigments are embedded in this membrane.
• Bidlack, J.E.; Stern, K.R.; Jansky, S. (2003).  Introductory Plant Biology . New York: McGraw-Hill. ISBN 978-0-07-290941-8.
• Blankenship, R.E. (2014).  Molecular Mechanisms of Photosynthesis  (2nd ed.). John Wiley & Sons. ISBN 978-1-4051-8975-0.
• Reece J.B., et al. (2013).  Campbell Biology . Benjamin Cummings. ISBN 978-0-321-77565-8.

Related Posts

8.1 Overview of Photosynthesis

Learning objectives.

In this section, you will explore the following questions:

• What is the relevance of photosynthesis to living organisms?
• What are the main cellular structures involved in photosynthesis?
• What are the substrates and products of photosynthesis?

Connection for AP ® Courses

As we learned in Chapter 7, all living organisms, from simple bacteria to complex plants and animals, require free energy to carry out cellular processes, such as growth and reproduction. Organisms use various strategies to capture, store, transform, and transfer free energy, including photosynthesis. Photosynthesis allows organisms to access enormous amounts of free energy from the sun and transform it to the chemical energy of sugars. Although all organisms carry out some form of cellular respiration, only certain organisms, called photoautotrophs, can perform photosynthesis. Examples of photoautotrophs include plants, algae, some unicellular eukaryotes, and cyanobacteria. They require the presence of chlorophyll, a specialized pigment that absorbs certain wavelengths of the visible light spectrum to harness free energy from the sun. Photosynthesis is a process where components of water and carbon dioxide are used to assemble carbohydrate molecules and where oxygen waste products are released into the atmosphere. In eukaryotes, the reactions of photosynthesis occur in chloroplasts; in prokaryotes, such as cyanobacteria, the reactions are less localized and occur within membranes and in the cytoplasm. (The structural features of the chloroplast that participate in photosynthesis will be explored in more detail later in The Light-Dependent Reactions of Photosynthesis and Using Light Energy to Make Organic Molecules.) Although photosynthesis and cellular respiration evolved as independent processes—with photosynthesis creating an oxidizing atmosphere early in Earth’s history—today they are interdependent. As we studied in Cellular Respiration, aerobic cellular respiration taps into the oxidizing ability of oxygen to synthesize the organic compounds that are used to power cellular processes.

Information presented and the examples highlighted in the section support concepts and learning objectives outlined in Big Idea 1 and Big Idea 2 of the AP ® Biology Curriculum Framework, as shown in the table. The learning objectives listed in the Curriculum Framework provide a transparent foundation for the AP ® Biology course, an inquiry-based laboratory experience, instructional activities, and AP ® exam questions. A learning objective merges required content with one or more of the seven science practices.

Teacher Support

Use this first part of the chapter to present an overview that will be filled out and completed in the later two portions. This will introduce the students to the biochemistry that they need to know and give them a chance to build up their understanding of the material.

Importance of Photosynthesis

Use this section to stress the importance of the interdependence between different species and the role played by photosynthesis in bringing energy to the living organisms. A number of terms, such as photoautotroph, heterotrophy, and chemoautotroph will be introduced here.

Photosynthesis is essential to all life on earth; both plants and animals depend on it. It is the only biological process that can capture energy that originates in outer space (sunlight) and convert it into chemical compounds (carbohydrates) that every organism uses to power its metabolism. In brief, the energy of sunlight is captured and used to energize electrons, whose energy is then stored in the covalent bonds of sugar molecules. How long lasting and stable are those covalent bonds? The energy extracted today by the burning of coal and petroleum products represents sunlight energy captured and stored by photosynthesis almost 200 million years ago.

Plants, algae, and a group of bacteria called cyanobacteria are the only organisms capable of performing photosynthesis ( Figure 8.2 ). Because they use light to manufacture their own food, they are called photoautotrophs (literally, “self-feeders using light”). Other organisms, such as animals, fungi, and most other bacteria, are termed heterotrophs (“other feeders”), because they must rely on the sugars produced by photosynthetic organisms for their energy needs. A third very interesting group of bacteria synthesize sugars, not by using sunlight’s energy, but by extracting energy from inorganic chemical compounds; hence, they are referred to as chemoautotrophs .

The importance of photosynthesis is not just that it can capture sunlight’s energy. A lizard sunning itself on a cold day can use the sun’s energy to warm up. Photosynthesis is vital because it evolved as a way to store the energy in solar radiation (the “photo-” part) as energy in the carbon-carbon bonds of carbohydrate molecules (the “-synthesis” part). Those carbohydrates are the energy source that heterotrophs use to power the synthesis of ATP via respiration. Therefore, photosynthesis powers 99 percent of Earth’s ecosystems. When a top predator, such as a wolf, preys on a deer ( Figure 8.3 ), the wolf is at the end of an energy path that went from nuclear reactions on the surface of the sun, to light, to photosynthesis, to vegetation, to deer, and finally to wolf.

Science Practice Connection for AP® Courses

Think about it.

• Why do scientists think that photosynthesis evolved before aerobic cellular respiration?
• Why do carnivores, such as lions, depend on photosynthesis to survive? What evidence supports the claim that photosynthesis and cellular respiration are interdependent processes?
• The first Think About It question is an application of Learning Objective 1.15 and Science Practice 7.2 because students are describing the evolution of two energy-procuring processes that today are present in different organisms.
• The second Think About It question is an application of Learning Objective 2.5 and Science Practice 6.2 because you are explaining how the interdependent processes of photosynthesis and cellular respiration allow organisms to capture, store, and use free energy.

Possible answers:

• Aerobic cellular respiration requires free oxygen, which was not available in the Earth’s atmosphere until photosynthetic organisms produced enough oxygen as waste to support developing aerobic respiration.
• Carnivores at the top of the food chain eat herbivores that eat photoautotrophs. So no matter where you are in the food chain, every species depends on photosynthesis to convert light energy to chemical energy. In ecosystems that lack photosynthetic organisms (such as by forests burned by forest fire), organisms on all levels of the food chain die off.

The structures, substrates and products of photosynthesis are introduced in this section. Remind them that Figure 8.5 can also be read from right to left, if cellular respiration is the subject. This should help the students to connect the two pathways of photosynthesis and cellular respiration.

Obtain diagrams of leaf structures to illustrate the content of this section. Try to bring in some leaves for students to look at. They have all seen lots of leaves, but probably never examined them for structural detail. A simple magnifying glass should allow them to see the inner structures discussed in this section.

Main Structures and Summary of Photosynthesis

Photosynthesis is a multi-step process that requires sunlight, carbon dioxide (which is low in energy), and water as substrates ( Figure 8.4 ). After the process is complete, it releases oxygen and produces glyceraldehyde-3-phosphate (G3P), simple carbohydrate molecules (which are high in energy) that can subsequently be converted into glucose, sucrose, or any of dozens of other sugar molecules. These sugar molecules contain energy and the energized carbon that all living things need to survive.

The following is the chemical equation for photosynthesis ( Figure 8.5 ):

Although the equation looks simple, the many steps that take place during photosynthesis are actually quite complex. Before learning the details of how photoautotrophs turn sunlight into food, it is important to become familiar with the structures involved.

In plants, photosynthesis generally takes place in leaves, which consist of several layers of cells. The process of photosynthesis occurs in a middle layer called the mesophyll . The gas exchange of carbon dioxide and oxygen occurs through small, regulated openings called stomata (singular: stoma), which also play roles in the regulation of gas exchange and water balance. The stomata are typically located on the underside of the leaf, which helps to minimize water loss. Each stoma is flanked by guard cells that regulate the opening and closing of the stomata by swelling or shrinking in response to osmotic changes.

In all autotrophic eukaryotes, photosynthesis takes place inside an organelle called a chloroplast . For plants, chloroplast-containing cells exist in the mesophyll. Chloroplasts have a double membrane envelope (composed of an outer membrane and an inner membrane). Within the chloroplast are stacked, disc-shaped structures called thylakoids . Embedded in the thylakoid membrane is chlorophyll, a pigment (molecule that absorbs light) responsible for the initial interaction between light and plant material, and numerous proteins that make up the electron transport chain. The thylakoid membrane encloses an internal space called the thylakoid lumen . As shown in Figure 8.6 , a stack of thylakoids is called a granum , and the liquid-filled space surrounding the granum is called stroma or “bed” (not to be confused with stoma or “mouth,” an opening on the leaf epidermis).

Visual Connection

• Rate of photosynthesis will be inhibited as the level of carbon dioxide decreases.
• Rate of photosynthesis will be inhibited as the level of oxygen decreases.
• The rate of photosynthesis will increase as the level of carbon dioxide increases.
• Rate of photosynthesis will increase as the level of oxygen increases.

The Two Parts of Photosynthesis

There are different terms that have been used for these reactions. Go over each pair of terms and discuss how they apply to the pathways.

Photosynthesis takes place in two sequential stages: the light-dependent reactions and the light independent-reactions. In the light-dependent reactions , energy from sunlight is absorbed by chlorophyll and that energy is converted into stored chemical energy. In the light-independent reactions , the chemical energy harvested during the light-dependent reactions drives the assembly of sugar molecules from carbon dioxide. Therefore, although the light-independent reactions do not use light as a reactant, they require the products of the light-dependent reactions to function. In addition, several enzymes of the light-independent reactions are activated by light. The light-dependent reactions utilize certain molecules to temporarily store the energy: These are referred to as energy carriers. The energy carriers that move energy from light-dependent reactions to light-independent reactions can be thought of as “full” because they are rich in energy. After the energy is released, the “empty” energy carriers return to the light-dependent reaction to obtain more energy. Figure 8.7 illustrates the components inside the chloroplast where the light-dependent and light-independent reactions take place.

Link to Learning

Click the link to learn more about photosynthesis.

• The light reactions produces ATP and NADPH, which are then used in the Calvin cycle.
• The light reactions produces NADP + and ADP, which are then used in the Calvin cycle.
• The light reactions uses NADPH and ATP, which are produced by the Calvin cycle.
• The light reactions produce only NADPH, which is produced by the Calvin cycle.

Everyday Connection for AP® Courses

Photosynthesis at the grocery store.

Major grocery stores in the United States are organized into departments, such as dairy, meats, produce, bread, cereals, and so forth. Each aisle ( Figure 8.8 ) contains hundreds, if not thousands, of different products for customers to buy and consume.

Although there is a large variety, each item links back to photosynthesis. Meats and dairy link, because the animals were fed plant-based foods. The breads, cereals, and pastas come largely from starchy grains, which are the seeds of photosynthesis-dependent plants. What about desserts and drinks? All of these products contain sugar—sucrose is a plant product, a disaccharide, a carbohydrate molecule, which is built directly from photosynthesis. Moreover, many items are less obviously derived from plants: For instance, paper goods are generally plant products, and many plastics (abundant as products and packaging) are derived from algae. Virtually every spice and flavoring in the spice aisle was produced by a plant as a leaf, root, bark, flower, fruit, or stem. Ultimately, photosynthesis connects to every meal and every food a person consumes.

• at the base
• near the top
• in the middle, but generally closer to the top
• in the middle, but generally closer to the base

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• Authors: Julianne Zedalis, John Eggebrecht
• Publisher/website: OpenStax
• Book title: Biology for AP® Courses
• Publication date: Mar 8, 2018
• Location: Houston, Texas
• Book URL: https://openstax.org/books/biology-ap-courses/pages/1-introduction
• Section URL: https://openstax.org/books/biology-ap-courses/pages/8-1-overview-of-photosynthesis

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